UnsatisfiedLinkError when manually registering functions via JNI_OnLoad

I have an Android application which uses some JNI to communicate between my Java and C++ classes. The JNI_OnLoad function looks something like this:

JNIEXPORT jint JNI_OnLoad(JavaVM *vm, void */* reserved */)
{
  qDebug() << "starting to register functions";
  JNIEnv *env;
  if (vm->GetEnv(reinterpret_cast<void **>(&env), JNI_VERSION_1_6) != JNI_OK)
  {
    return JNI_ERR;
  }
  jclass nativeClass = env->FindClass("com/example/myapp/MainActivity");
  if (!nativeClass) return JNI_ERR;
  if (env->RegisterNatives(nativeClass, mainActivityMethods, sizeof(mainActivityMethods) / sizeof(mainActivityMethods[0])) < 0)
  {
    return JNI_ERR;
  }
  qDebug() << "registered all functions!";
  return JNI_VERSION_1_6;
}

It is contained in a file which is definitely listed as a source file in the .pro file. However, the debug statements are never printed out and I get

java.lang.UnsatisfiedLinkError: No implementation found for java.lang.String 
com.example.myapp.MainActivity.foo() 
(tried Java_com_example_myapp_MainActivity_foo
and Java_com_example_myapp_MainActivity_foo__)

I did not use to get this error before I upgraded to 5.11. Everything was working fine previously.

It is, however, worth noting that I also did a "VCS refresh" in the sense that I had to pull all the files from the repository into a fresh directory because the old one was deleted. This could potentially be a factor, though I don't think so.

I was able to manually resolve the error by adding a call to `System.loadLibrary("MyApp")` However, I was under the impression this was not supposed to be necessary. Therefore I believe it is a bug.